A PhD program in Biology is a vast and profoundly dynamic and research-intensive discipline dedicated to the systematic study of living organisms, their structure, function, growth, evolution, distribution, and the intricate molecular, cellular, and ecological processes that sustain and define life in all its extraordinary diversity across the planet. Rooted in cell biology, molecular genetics, biochemistry, physiology, ecology, evolutionary biology, and bioinformatics, the program develops scientists with the theoretical depth, experimental rigor, and analytical sophistication to investigate biological phenomena across scales ranging from the molecular architecture of the cell to the complex dynamics of ecosystems and the evolutionary history of life on Earth.